Pressure chamber is to stablise tank pressure - ie pressure into the engine. Some people will run very long fuel line at the same time.... Think of it as a big air cushion. If you can tell the difference, you are a very good RC player!!
I need to dig out my physics text book - since when the location of an 'air' chamber will affect pressure. I think it is the length/diameter of the fuel line that affects pressure, the longer the line, the more pressure drop and smaller diameter, higher pressure?